Great Britain's Jody Cundy saw his 20-year reign as para cycling's C4 kilo time trial world champion come to an end at the para cycling track world championships in the Netherlands.
The 48-year-old, who won his first world title in 2006 and had held the rainbow jersey through 15 consecutive championships, finished with silver after completing the kilometre in one minute and 4.93 seconds.
Australian Korey Boddington claimed gold, beating Cundy's time by 2.25 seconds on the final run.
Before the event, Cundy acknowledged that "age was catching up with him" and that adding to his 23 track cycling world titles would be "hard."
Cundy is a nine-time Paralympic champion, with three of his medals coming in para swimming between 1996 and 2000.
Elsewhere on day one of the championships, Great Britain's Rebecca Newark won gold in the women's C5 kilo time trial.
In the women's B kilo time trial, Britain's Sophie Unwin and pilot Jenny Holl took silver, while Elizabeth Jordan and pilot Anna Whitworth Hay claimed bronze.
British pair Morgan Fice-Noyes and Christopher Latham earned silver in the men's 4km pursuit, and Finlay Graham added bronze in the men's C3 sprint final.
